As a taxonomist, he circumscribed a number of species within the grass genus " Puccinellia ".
One of these, " Puccinellia fasiculata ", commonly " Borrer's saltmarsh grass, " is protected, along with its habitat.
In early winter the wigeon select " Puccinellia maritima " in preference to " Agrostis stolonifera " and " Festuca rubra ".
There are several exclusive species in the national park, including, " Puccinellia svalbardensis ", " Gentianella tenella " and " Kobresia simpliciuscula ".
The area of ditch close to Breydon Water are more brackish in nature and feature species such as soft hornwort " Ceratophyllum submersum " and the nationally scarce stiff saltmarsh grass " Puccinellia rupestris ".
Adjacent to these saline flashes are areas of saltmarsh vegetation containing species such as sea aster " Aster tripolium ", lesser sea-spurrey " Spergularia marina " and reflexed saltmarsh-grass " Puccinellia distans ".
A distinct habitat is provided by the interface between saltmarsh and dune and supports reflexed saltmarsh grass ( " Puccinellia distans " ), hard-grass ( " Parapholis strigosa " ) and sea-milkwort ( " Glaux maritima " ).
The best remaining saltmarsh is at the eastern, seaward end of the reedbeds and these support locally rare saltmarsh plants including sea club-rush " Bolboschoenus maritimus ", grey bulrush " Schoenoplectus tabernaemontani " and common saltmarsh grass " Puccinellia maritima ".