The development process , from fertilized egg to spiracular formation can be divided into 26 stages , took 212 . 94 hours under the water temperature of ( 20 0 . 5 ) and 170 . 95 hours under the natural temperature ( 21 . 5 24 ) . the external morphological characteristics in various stages of the embryonic development were similar to those of rana nigromaculata , but the hatching stage was later than that of rana nigromaculata and earlier than that of rana limnocharis 自受精卵期至鳃盖完成期共分为26个时期，其发育历程及各时期胚胎外形特征与黑斑蛙基本相似，但孵化期比黑斑蛙推迟三个胚期而早于泽蛙，在水温( 20 0 . 5 )及常温（ 21 . 5 24 ）条件下，其胚胎发育的全时程分别为212 . 94小时和170 . 95小时。
It requires further investigation whether spermatogonial proliferation starts in november . 3 . interstitial cells synthesize t in spring and autumn , its spermatogonial proliferation independent on the levels of t . t in the follicular cells in the testis of rana quadranus regulate sperm maturation by paracrine mechanism 隆肛蛙因对寒冷的气候很敏感，在外界活动的时间短，使其从精原细胞到成熟精子发育的时间很短，精原细胞很快增殖并进行成熟分裂而成为成熟精子。
